Also Ref. 90102. Recorded from Meiro R., Madang, Umboi Is. (Ref. 6192), Kiriwina Is. and the Maiwara R. mouth of west of Alotau (Ref. 6771). Occurs in estuarine and freshwater habitats, frequently where mangroves are abundant; in coastal creeks, it is usually found within 2-3 km of the sea (Ref. 2847).